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- Francisco Ayala + (a Spanish-American evolutionary biologist … a Spanish-American evolutionary biologist and philosopher at the University of California, Irvine.[2] He is a former Dominican priest,[3][4] ordained in 1960,[5] but left the priesthood that same year. After graduating from the University of Salamanca, he moved to the United States in 1961 to study for a PhD at Columbia University. There, he studied for his doctorate under Theodosius Dobzhansky, graduating in 1964.[6] He became a US citizen in 1971.n 1964.[6] He became a US citizen in 1971.)